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Constants — the measured numbers

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Every load-bearing constant with its units, scope, evidence class, source and falsifier.

307 rows.

name symbol value units scope class source falsifier
A4 A4 440 ± 0.5 Hz Western concert pitch, by convention, at ~20 °C OBSERVED-REPLICATED (as a standard, not a natural constant) ISO 16:1975, Acoustics — Standard tuning frequency It is a convention; only the claim that it is natural is falsifiable, and that claim is inadmissible.
T_ant T_ant ≈3.5 K (excess antenna temperature at 4080 Mc/s) Holmdel horn antenna, 1964–65; survived removal of every known instrumental/atmospheric term, incl. the pigeons OBSERVED-REPLICATED Penzias & Wilson (1965), ApJ 142:419–421 An instrumental or local source reproducing the excess.
T_CMB T_CMB 2.72548 ± 0.00057 K Present-day CMB monopole temperature OBSERVED-REPLICATED Fixsen (2009), ApJ 707:916 A measurement outside the stated interval by a calibrated instrument.
SI defining constants SI defining constants Δν_Cs = 9 192 631 770 Hz; c = 299 792 458 m/s; h = 6.626 070 15 × 10⁻³⁴ J·s; e = 1.602 176 634 × 10⁻¹⁹ C; k = 1.380 649 × 10⁻²³ J/K; N_A = 6.022 140 76 × 10²³ mol⁻¹ Hz, m/s, J·s, C, J/K, mol⁻¹ exact by convention, effective 20 May 2019 (NO evidence class — chapter states: 'Definitional values, deliberately carrying NO evidence class … They are not observations of nature and this wing's evidence classes do not apply to them.') BIPM, SI Brochure, 9th ed., 2019 Not falsifiable — conventions, not observations. The chapter's instruction is: know which of your numbers are conventions.
N_rod N_rod 92 × 10⁶ (range 77.9–107.3 × 10⁶) cells human retina; 8 wholemounts, 7 donors, ages 27–44 OBSERVED-REPLICATED Curcio, Sloan, Kalina & Hendrickson (1990), J Comp Neurol 292:497–523, DOI 10.1002/cne.902920402 Recount in a comparable cohort; a mean outside the stated range refutes.
N_cone N_cone 4.6 × 10⁶ (range 4.08–5.29 × 10⁶) cells as above OBSERVED-REPLICATED Curcio et al. (1990) As above.
D_cone,fovea D_cone,fovea 199,000 (range 100,000–324,000) cones/mm² human foveal peak; same cohort OBSERVED-REPLICATED Curcio et al. (1990) As above.
N_optic N_optic 1,159,000 ± 196,000 (range 816,000–1,502,000) axons human optic nerve; 22 nerves, 19 subjects, ages 20–75 OBSERVED-REPLICATED Jonas, Müller-Bergh, Schlötzer-Schrehardt & Naumann (1990), Invest Ophthalmol Vis Sci 31(4):736–744 Recount; a mean outside the stated range refutes.
r_retina r_retina ≈ 83 : 1 dimensionless human visual blanket, order-of-magnitude only MODELED — assumptions: (N_rod + N_cone)/N_optic across different cohorts, unpaired, no per-eye matching, ignores non-uniform convergence (foveal ≈ 1:1 vs peripheral ≫ 100:1) arithmetic on Curcio et al. (1990) + Jonas et al. (1990) Measure both counts in the same eyes. A paired ratio outside ~50–150:1 refutes this estimate.
f_aff (rat) f_aff (rat) ~80% afferent / 20% efferent % of fibers rat, abdominal vagus OBSERVED-CONTESTED — the widely-quoted '80% of the vagus is afferent', routinely cited outside this scope Prechtl & Powley (1990), Anat Embryol 181:101–115, DOI 10.1007/BF00198950 Measure the human cervical vagus and obtain 80% ± small. Kronsteiner et al. (2024) did, and did not.
f_aff (human) f_aff (human) sensory 73.9 ± 7.5% (R), 72.4 ± 5.6% (L); parasympathetic 13.2 ± 1.8% / 13.3 ± 3.0%; sympathetic 13 ± 5.9% / 14.3 ± 4.0% % of fibers human, cervical vagus; 8 cadavers, immunofluorescence OBSERVED-CONTESTED — carry with the row above; both positions stand Kronsteiner et al. (2024), Brain Stimulation 17(3):510–524, DOI 10.1016/j.brs.2024.04.016 Independent replication in a larger cohort; a sensory fraction outside ~65–82% refutes.
N_vagus N_vagus ~100,000 (light microscopy, 1961) vs 25,489 ± 2,781 (R) / 23,286 ± 3,164 (L) (modern, 2024) axons human cervical vagus OBSERVED-CONTESTED — a ~4× disagreement between methods. The dispute is the finding; the modern claim is that light microscopy cannot resolve unmyelinated fibers Hoffman & Schnitzlein (1961), Anat Rec 139(3), DOI 10.1002/ar.1091390312; Kronsteiner et al. (2024) Blinded EM recount across labs on shared specimens. Convergence on either value resolves it.
ν (water) ν (water) ~10⁻² cm² s⁻¹ liquid water, room temp OBSERVED-REPLICATED Purcell (1977) Standard viscometry.
Capillary length ℓ_c Capillary length ℓ_c ≈2.6 mm air–water, σ ≈ 0.07 N m⁻¹ OBSERVED-REPLICATED Bush & Hu (2006) Direct meniscus measurement.
Min. capillary wave speed c_m Min. capillary wave speed c_m 23 cm s⁻¹ air–water interface OBSERVED-REPLICATED Lighthill (1979), via Bush & Hu (2006) Waves radiated by steady motion below 23 cm s⁻¹.
λ (air) λ (air) ≈68 nm 1 atm, 25 °C OBSERVED-REPLICATED standard kinetic theory Direct mean-free-path measurement.
Earth mantle viscosity Earth mantle viscosity 10²¹ poise mantle flow MODELED (geophysical inference) quoted in Purcell (1977) Independent rheological determination.
Pr Pr 7 (water) / 0.71 (air) / 0.025 (mercury) near room temperature OBSERVED-REPLICATED standard property tables Property measurement.
D_GFP,ec D_GFP,ec 7.7 ± 2.5 µm²/s GFP (27 kDa), E. coli cytoplasm, FRAP/photoactivation OBSERVED-REPLICATED BNID 100193; Elowitz et al. 1999, J Bacteriol 181(1):197–203 Repeat FRAP; a value outside 3–14 µm²/s under stated conditions refutes.
D_GFP,euk D_GFP,euk 27 µm²/s GFP-S65T, CHO cytoplasm OBSERVED-REPLICATED BNID 101997; Swaminathan et al. 1997, Biophys J 72(4):1900–7 Independent FRAP in eukaryotic cytoplasm disagreeing >2×.
L_Thio L_Thio 100–300 (max 750) µm Thiomargarita namibiensis, cell width OBSERVED-REPLICATED Schulz et al. 1999, Science 284:493–495 Larger true-cytoplasm cell found.
f_vac f_vac 80–98 % of cell volume Thiomargarita nitrate vacuole; cytoplasm shell ~1–2 µm OBSERVED-REPLICATED Schulz et al. 1999 Show cytoplasm fills the cell.
ATP_ec ATP_ec ~10⁷ ATP/s/cell E. coli, growing OBSERVED-REPLICATED BNID 111461, 110656, 110628 Independent measurement >10× off.
ATP_mam ATP_mam ~10⁹ ATP/s/cell human fibroblast, ~3,000 µm³ OBSERVED-REPLICATED BNID 111476 As above.
P_ec P_ec ~10⁻¹² (1,000 W/kg) W/cell E. coli, glucose minimal media OBSERVED-REPLICATED BNID 109687 As above.
c_pep c_pep 4 ATP per peptide bond 2 (PPi, aa-tRNA charging) + 1 GTP × 2 elongation factors OBSERVED-REPLICATED Milo & Phillips 2015, Cell Biology by the Numbers Show a bond formed for <4.
ΔG_ATP ΔG_ATP −47 to −50 (≈20 k_BT ≈ 80–90 pN·nm) kJ/mol in vivo; E. coli on glucose −47 OBSERVED-REPLICATED BioNumbers (How much energy is released in ATP hydrolysis?) Measured phosphorylation potential outside −40 to −65.
ΔG°'_ATP ΔG°'_ATP −28 to −34 (≈12 k_BT) kJ/mol standard conditions (1 M) — NOT the cell OBSERVED-REPLICATED as above
τ_F1 τ_F1 ~40 pN·nm F₁-ATPase torque, constant across load/speed OBSERVED-REPLICATED Yasuda et al. 1998, Cell 93:1117–1124 Load-dependent torque under same assay.
W_F1 W_F1 ~80 (vs ~90 available) pN·nm per 120° step F₁, single-molecule, in vitro OBSERVED-REPLICATED Yasuda et al. 1998; Noji et al. 1997, Nature 386:299–302 Work/step measured well below 80.
d_kin d_kin 8 nm/step kinesin-1 on microtubule, optical trap OBSERVED-REPLICATED Svoboda et al. 1993, Nature 365:721–727 A different step periodicity.
n_ATP,kin n_ATP,kin 1 ATP per 8-nm step kinesin-1 OBSERVED-REPLICATED Schnitzer & Block 1997, Nature 388:386–390 Measured coupling ≠ 1:1.
F_stall,kin F_stall,kin 5–6 or 7–8 pN 5–6: Svoboda & Block 1994. 7–8: force clamp, Visscher 1999 OBSERVED-CONTESTED Svoboda & Block 1994, Cell 77:773–784; Visscher et al. 1999, Nature 400:184–189 Resolve by assay; do not average. A study reconciling both under one method.
v_kin v_kin ~0.5–1 (commonly ~0.8) µm/s saturating ATP, near-zero load, in vitro OBSERVED-REPLICATED Svoboda & Block 1994 (force–velocity) Outside range under stated conditions.
d_myoV d_myoV ~36 nm/step myosin-V on actin; stall ~2–3 pN OBSERVED-REPLICATED single-molecule optical trap literature Different step periodicity.
d_dyn d_dyn 8 (or 8→32, load-dependent) nm/step cytoplasmic dynein — unresolved OBSERVED-CONTESTED Optical-tweezer reports disagree A method resolving load-dependence.
ε_rib ε_rib 10⁻⁴–10⁻³ per codon missense/misreading OBSERVED-REPLICATED Kramer & Farabaugh 2007, RNA 13:87–96 Measured rate outside range.
ε_pol ε_pol ~10⁻⁴–10⁻⁵ per nt polymerase base selectivity ALONE OBSERVED-REPLICATED Kunkel & Bebenek 2000, Annu Rev Biochem; Kunkel 2004, JBC Exonuclease-deficient rate outside range.
ε_proof ε_proof ~10⁻⁶–10⁻⁷ (×10²–10³ gain) per nt + exonucleolytic proofreading OBSERVED-REPLICATED as above MMR-deficient rate outside range.
ε_final ε_final 10⁻⁸–10⁻¹⁰ per nt + mismatch repair; pro- and eukaryotes OBSERVED-REPLICATED as above Whole-genome mutation accumulation outside range.
d_bilayer d_bilayer 4–5 nm lipid bilayer thickness OBSERVED-REPLICATED standard membrane biophysics; Milo & Phillips 2015 Structural measurement outside range.
V_m V_m ~−70 mV resting neuron OBSERVED-REPLICATED standard electrophysiology
E_m E_m 1.4–1.8 × 10⁷ V/m V/d, 70 mV over 4–5 nm MODELED Computed in-chapter from V_m and d_bilayer Arithmetic error, or d/V refuted.
E_air E_air ≈3 × 10⁶ V/m dry air, 1 atm — dielectric strength OBSERVED-REPLICATED (not primary-sourced in this pass) standard physical reference Fetch a primary reference.
C_m C_m ~1 (0.01) µF/cm² (F/m²) specific membrane capacitance, near-invariant across cell types OBSERVED-REPLICATED standard membrane biophysics (Cole; Hodgkin & Huxley 1952) A cell type deviating >2× with intact bilayer.
κ_MT κ_MT 2.2 × 10⁻²³ (±6.4%); 2.1 × 10⁻²³ (±4.7%, rhodamine) N·m² taxol-stabilised microtubule, flexural rigidity OBSERVED-REPLICATED Gittes et al. 1993, J Cell Biol 120(4):923–934 Independent measurement >2× off.
ℓ_p,MT ℓ_p,MT ~5,200 (5.2 mm) µm microtubule persistence length, ℓ_p = κ/k_BT OBSERVED-REPLICATED Gittes et al. 1993 see contested row below.
ℓ_p,actin ℓ_p,actin ~17.7 µm actin filament, rhodamine-phalloidin OBSERVED-REPLICATED Gittes et al. 1993 Independent measurement >2× off.
n_Na/ATP n_Na/ATP 3 Na⁺ per 1 ATP ions/ATP Na⁺/K⁺-ATPase stoichiometry OBSERVED-REPLICATED Attwell & Laughlin 2001 and standard references Measured stoichiometry ≠ 3:2:1.
E_Schumann@membrane E_Schumann@membrane V/m field amplitude at a cell membrane NOT-MEASURED not sourced in this pass Measure amplitude; compare to k_BT and membrane noise.
λ_Bcd λ_Bcd ≈ 100 μm D. melanogaster early embryo, L ≈ 490 μm OBSERVED-REPLICATED Gregor et al. (2007a), Cell 130(1):141–152 Independent Bcd-GFP profiling giving a decay constant far outside ~80–120 μm at standard temperature.
L L ≈ 490 μm same embryo, AP axis OBSERVED-REPLICATED Gregor et al. (2007a) Direct imaging outside ~450–550 μm.
D_Bcd D_Bcd 0.30 ± 0.09 (FRAP); 0.37 ± 0.05 (indirect) μm²/s cortical cytoplasm, cycles 10–14 OBSERVED-CONTESTED — incompatible with λ ≈ 100 μm under SDD Gregor et al. (2007a); dispute printed in Grimm et al. (2010), Development 137(14):2253–2264 A method giving D large enough that √(Dτ) reproduces λ within the ~1 h formation window would dissolve the tension.
Δc/c Δc/c ≈ 10 % adjacent nuclei (~8 μm apart) at the hb boundary, cycle 14 OBSERVED-REPLICATED Gregor et al. (2007b), Cell 130(1):153–164 Measured inter-nuclear Bcd difference at the boundary ≫ or ≪ 10%.
σ_x (hb domain) σ_x (hb domain) 2–3 % egg length hb transcription domain position, embryo-to-embryo OBSERVED-REPLICATED Gregor et al. (2007b) Reproducibility measured far worse (≫3% EL) in a clean prep.
σ_x (4 gap genes) σ_x (4 gap genes) ≈ 1 % egg length joint gap-gene readout, AP axis, near-constant along axis OBSERVED-REPLICATED Dubuis et al. (2013), PNAS 110(41):16301–16308 Decoding a fresh dataset yielding error ≫1% EL.
σ (limb bud mesoderm) σ (limb bud mesoderm) 20.1 dyn/cm chick embryonic tissue aggregate, parallel-plate compression OBSERVED-REPLICATED Foty et al. (1996), Development 122(5):1611–1620 Remeasurement inverting the envelopment hierarchy.
σ (pigmented epithelium / heart / liver / neural retina) σ (pigmented epithelium / heart / liver / neural retina) 12.6 / 8.5 / 4.6 / 1.6 dyn/cm as above OBSERVED-REPLICATED Foty et al. (1996) A tissue enveloping one of higher measured σ.
r_p r_p 8.4075(64) × 10⁻¹⁶ m proton rms charge radius OBSERVED-REPLICATED CODATA 2022 (NIST) Next CODATA adjustment moves it beyond stated u.
α_s(m_Z) α_s(m_Z) 0.1180 ± 0.0009 dimensionless strong coupling at the Z mass; runs to O(1) at ~1 GeV OBSERVED-REPLICATED PDG world average (recent editions give 0.1179–0.1180 ± 0.0009) A PDG edition outside 0.117–0.119.
a₀ a₀ 5.291 772 105 44(82) × 10⁻¹¹ m Bohr radius OBSERVED-REPLICATED CODATA 2022 (NIST) Next CODATA adjustment beyond stated u.
α⁻¹ α⁻¹ 137.035 999 177(21) dimensionless fine-structure constant; α = 7.297 352 5643(11) × 10⁻³ OBSERVED-REPLICATED CODATA 2022 (NIST) As above.
E_ion,H E_ion,H 13.6 eV hydrogen ionization OBSERVED-REPLICATED (standard reference; primary not read in this pass) standard atomic physics (NIST ASD) Fetch NIST ASD; a value outside 13.59–13.60.
E_C–C E_C–C ~348 kJ/mol C–C bond dissociation enthalpy OBSERVED-REPLICATED (standard reference; primary not read in this pass) standard thermochemical tables Fetch a primary table; value outside ~330–360.
ΔG_fold ΔG_fold ~20–63 (5–15 kcal/mol) kJ/mol net protein folding stability, typical globular OBSERVED-REPLICATED (standard range; primary not sourced in this pass) standard protein biophysics Locate a primary survey; range refuted.
ν_water ν_water ~1.0 × 10⁻⁶ m²/s kinematic viscosity, 20 °C OBSERVED-REPLICATED (standard reference) standard fluid-property tables Measurement outside ~0.9–1.1 × 10⁻⁶ at 20 °C.
R⊙ R⊙ 6.957 × 10⁸ m IAU nominal solar radius (exact by adoption, not a CBE) OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016, AJ 152:41 IAU re-adoption.
L⊙ L⊙ 3.828 × 10²⁶ W IAU nominal solar luminosity OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016 As above.
S⊙ S⊙ 1361 W/m² IAU nominal total solar irradiance OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016 As above.
T_eff,⊙ T_eff,⊙ 5772 K IAU nominal solar effective temperature OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016 As above.
(GM)⊙ (GM)⊙ 1.3271244 × 10²⁰ m³/s² IAU nominal solar mass parameter OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016 As above.
R_eE R_eE 6.3781 × 10⁶ m IAU nominal terrestrial equatorial radius (polar: 6.3568 × 10⁶) OBSERVED-REPLICATED (adopted constant) IAU 2015 Res. B3; Prša et al. 2016 As above.
AU AU 1.495 978 707 × 10¹¹ m astronomical unit, exact by IAU definition OBSERVED-REPLICATED (defined) IAU 2012 Res. B2 (definition; primary not read in this pass) Fetch IAU 2012 Res. B2.
D25_MW D25_MW 26.8 ± 1.1 (≈8.3 × 10²⁰ m) kpc Milky Way isophotal diameter OBSERVED-CONTESTED (secondary summary; primary not read in this pass) attributed to Goodwin et al. 1997/98, The Observatory 118:201–208 Read the primary; an independent estimate outside 25.7–27.9 kpc.
R₀ R₀ 8178 ± 13(stat) ± 22(sys) (≈ 8.178 kpc; 0.16% stat, 0.27% total) pc Sun → Galactic Centre; direct geometric measurement — S2's orbit via VLTI/GRAVITY interferometry + 27 yr astrometry/spectroscopy OBSERVED-REPLICATED (current standard reference) GRAVITY Collab. (Abuter et al.) 2019, A&A 625:L10, DOI 10.1051/0004-6361/201935656 (primary read in this pass) An independent geometric measurement outside ~8.13–8.23 kpc.
P/m E. coli P/m E. coli ~10³ (order-of-magnitude; wet-mass basis — see falsifier) W/kg E. coli strain C-3000, glucose minimal medium; a conversion, not a W/kg measurement MODELED Computed from BNID 109687 — an O₂ uptake rate, 30 mmol O₂/gDW/h — via the enthalpy of O₂ consumption (~478 kJ/mol O₂) ÷ a cell mass; chain via NA-08 Recomputed: 30 mmol/gDW/h × 478 kJ/mol ÷ 3600 s = ~4 × 10³ W/kg dry ≈ ~1.2 × 10³ W/kg wet at dry/wet ≈ 0.3 — the stated ~10³ closes on a wet basis only. NA-08's mass basis is NOT-CONFIRMED in this pass. Also: an independent measurement >10× off.
L⊙/M⊙ L⊙/M⊙ 1.9 × 10⁻⁴ W/kg Sun; 3.828 × 10²⁶ W ÷ 1.988 × 10³⁰ kg MODELED Computed in-chapter from IAU 2015 nominal L⊙ and (GM)⊙/G (CODATA 2022) Arithmetic error.
n_IMA n_IMA 6,200 valid species IMA-CNMNC list, January 2026 — a dated snapshot, not a constant OBSERVED-REPLICATED IMA-CNMNC, The New IMA List of Minerals (2026-01) Consult current CNMNC master list; a different count. This row is built to go stale — see r_IMA.
d_Si–O d_Si–O ~1.62 (range 1.55–1.72) Å Si–O, tetrahedral coordination, silicate structures OBSERVED-REPLICATED (primary not read this pass) Si–O bond-length literature (Brown & Gibbs, Am Mineral; Cruickshank's rule ~1.63 Å) Refine a silicate structure; mean Si–O outside 1.55–1.72.
θ_O–Si–O θ_O–Si–O 109.47 degrees ideal regular tetrahedron; real angles distort MODELED Geometry; Hückel-energy minimum at ideal angle (Am Mineral 57:1614) A regular-tetrahedron silicate at a different angle.
N_sg N_sg 230 (32 point groups, 14 Bravais lattices, 7 systems) space groups periodic order in 3-D Euclidean space — a closed formal result about the space, not a survey of rocks MODELED (closed formal result; deliberately not an empirical class — no mineral could refute it) Fedorov (1891); Schoenflies (1891); list corrected to 230 by 1892 Formal, not empirical: re-derive the enumeration — exhibit a 231st periodic group, or a duplicate among the 230.
Al₂SiO₅ triple pt Al₂SiO₅ triple pt both carried: ~501 °C, ~3.8 kbar (1971) and 504 ± 20 °C, 3.75 ± 0.25 kbar (1993) °C, kbar kyanite–andalusite–sillimanite invariant point; scoped to the Holdaway line of work — the wider aluminosilicate triple-point literature is not carried in this row OBSERVED-CONTESTED Holdaway (1971), Am J Sci 271:97–131; reevaluated Holdaway & Mukhopadhyay (1993), Am Mineral 78:298–315 Re-run the brackets; coordinates outside 504 ± 20 °C, 3.75 ± 0.25 kbar. Do not average the two — the 1971 point falls inside the 1993 brackets, which is agreement, not licence to merge.
H_qtz H_qtz 12.1 ± 1.1 (micro) / 14.5 ± 0.4 (DSI) GPa quartz (0001) OBSERVED-REPLICATED Whitney, Broz & Cook (2007), Am Mineral 92:281–288, Table 1 Independent indentation on (0001) outside range.
E*_qtz E*_qtz 117 ± 3 GPa quartz (0001), indentation modulus OBSERVED-REPLICATED Whitney et al. (2007), Table 1 As above.
K_IC,qtz K_IC,qtz 1.5 ± 0.3 MPa·m^½ quartz (0001) OBSERVED-REPLICATED Whitney et al. (2007) As above.
H_orth H_orth 6.9 ± 0.7 (micro) / 9.1 ± 0.6 (DSI) GPa orthoclase (101), Mohs 6 OBSERVED-REPLICATED Whitney et al. (2007), Table 1 As above.
H_per H_per 5.3 ± 1.0 (micro) / 9.4 ± 1.4 (DSI) GPa periclase MgO, Mohs 6–6.5 — polycrystalline synthetic reference material, not a single crystal. Softer than orthoclase by microhardness only; the DSI column preserves Mohs order for the pair (9.4 ± 1.4 vs 9.1 ± 0.6, overlapping) OBSERVED-REPLICATED (the values) / scope-mismatched for any rank-inversion claim — Mohs is a single-crystal scratch test Whitney et al. (2007), Table 1 Re-indent single-crystal MgO; the microhardness inversion disappears.
K_IC,per K_IC,per 3.9 ± 0.8 MPa·m^½ periclase — toughest of that set, and among the softest; polycrystalline synthetic OBSERVED-REPLICATED Whitney et al. (2007) As above.
H_ky H_ky Mohs 5–5.5 on (001) vs Mohs 7 on (100)/(010); DSI 14.8 ± 1.4 on (001); E* 186 ± 8 → 253 ± 19 GPa kyanite — one crystal, two Mohs numbers, face-dependent; and (001), the table's lowest Mohs entry, is harder by DSI than orthoclase (9.1) and periclase (9.4) — the clean rank inversion: single-crystal, no reference-sample caveat OBSERVED-REPLICATED Whitney et al. (2007), Table 1 Indent both faces and find isotropy; or find (001) softer than orthoclase by DSI.
t½ ²³⁸U t½ ²³⁸U (4.4683 ± 0.0024) × 10⁹ yr OBSERVED-REPLICATED Jaffey et al. (1971), Phys Rev C 4:1889–1906 Re-measure specific activity; outside stated uncertainty.
t½ ²³⁵U t½ ²³⁵U (7.0381 ± 0.0048) × 10⁸ yr ratio to ²³⁸U = 6.35× — the concordia's leverage OBSERVED-REPLICATED Jaffey et al. (1971) As above.
t½ ⁴⁰K t½ ⁴⁰K (1.2522 ± 0.0027) × 10⁹ yr ⁴⁰K half-life (coverage factor not read this pass — fetch the primary) OBSERVED-REPLICATED DDEP 2025 evaluation; Mougeot et al., Metrologia 63(1) (2026), doi:10.1088/1681-7575/ae3733 Supersedes 1.248 ± 0.003 × 10⁹ — re-evaluate.
⁴⁰K branches ⁴⁰K branches β⁻→⁴⁰Ca 89.56(7)%; EC→⁴⁰Ar total ≈10.44% = EC* 10.34(7)% (excited state, 1460.8 keV γ) + EC₀ 0.098% (ground state, no γ). 89.56 + 10.44 = 100.00 % K–Ar uses EC₀ + EC*, i.e. all radiogenic ⁴⁰Ar — not EC* alone. Quoting 10.34% as 'the EC branch' drops a channel and loses 0.10% of the decay OBSERVED-REPLICATED DDEP 2025 evaluation (Mougeot et al. 2026); EC₀ first observed by the KDK collaboration, Phys Rev Lett 131:052503 (2023): I(EC₀) = 0.098% ± 0.023 (stat) ± 0.010 (sys) Branches that fail to sum to 100%; or an EC₀ re-measurement outside the KDK uncertainty.
t½ ⁸⁷Rb t½ ⁸⁷Rb 49.61 ± 0.16 Ga λ₈₇ = (1.3972 ± 0.0045) × 10⁻¹¹ a⁻¹ OBSERVED-REPLICATED Villa, De Bièvre, Holden & Renne (2015), GCA 164:382–385 Independent determination outside stated uncertainty.
t½ ¹⁴⁷Sm t½ ¹⁴⁷Sm 106.25 ± 0.38 Ga λ₁₄₇ = (6.524 ± 0.024) × 10⁻¹² a⁻¹, k = 2 OBSERVED-REPLICATED Villa et al. (2020), GCA (IUPAC-IUGS recommendation) As above.
t½ ¹⁴⁶Sm t½ ¹⁴⁶Sm 92.0 ± 2.6 (k = 1) — the current determination Ma direct re-determination by mass spectrometry + α-counting; agrees with neither legacy value OBSERVED-REPLICATED (one dedicated determination; not yet independently repeated) Chiera, Sprung, Amelin, Dressler, Schumann & Talip (2024), Sci Rep, doi:10.1038/s41598-024-64104-6 Independent re-determination outside 92.0 ± 2.6.
t_CAI t_CAI 4567.30 ± 0.16 Ma oldest solar-system solids; U-corrected Pb-Pb OBSERVED-REPLICATED Connelly et al. (2012), Science 338:651–655 Independent Pb-Pb outside uncertainty.
t_zircon t_zircon 4404 ± 8 Ma Jack Hills detrital zircon; δ¹⁸O 7.4→5.0‰ OBSERVED-REPLICATED Wilde, Valley, Peck & Graham (2001), Nature 409:175–178 An older confirmed terrestrial grain; or refute the δ¹⁸O inference.
d_borehole d_borehole 12,262 m Kola SG-3 — deepest direct sample of the Earth by true vertical depth (longer directional wells exist by measured length); ~0.19% of the way to the centre OBSERVED-REPLICATED Popov, Pevzner, Pimenov & Romushkevich (1999), Tectonophysics 306(3–4):345–366 A deeper true-vertical borehole; or a re-survey of SG-3 outside 12,262 m.
K_qtz, G_qtz, ρ_qtz K_qtz, G_qtz, ρ_qtz GPa, g/cm³ single-crystal adiabatic bulk/shear moduli, quartz & olivine NOT-MEASURED (not sourced this pass) Fetch Bass (1995), AGU Ref. Shelf 2:45–63.
T_ρmax T_ρmax 3.983 °C liquid H₂O, 1 atm OBSERVED-REPLICATED Wikipedia Properties of water (compiling standard reference data) Densimetry finding max outside 3.9–4.1 °C at 1 atm.
ρ(3.983 °C) ρ(3.983 °C) 0.99997495(84) g/mL as above OBSERVED-REPLICATED as above Value outside stated uncertainty.
ρ(0 °C) ρ(0 °C) 0.99984283(84) g/mL liquid, 1 atm OBSERVED-REPLICATED as above As above.
ρ(25 °C) ρ(25 °C) 0.99704702(83) g/mL liquid, 1 atm OBSERVED-REPLICATED as above As above.
ρ_ice ρ_ice 0.9167 g/mL ice Ih, 0 °C OBSERVED-REPLICATED Wikipedia Properties of water Structural/densimetric measurement outside ±0.001.
k_ice k_ice ~2.2 (~2.2–2.3; rises as T falls) W/(m·K) ice Ih, 0 °C, 1 atm — higher than liquid water OBSERVED-REPLICATED (secondary compilation) Wikipedia List of thermal conductivities (CRC-sourced row, ~273 K); closure: fetch a primary Measurement at 0 °C, 1 atm outside ~2.1–2.4.
k_water k_water ~0.561 W/(m·K) liquid water, 0 °C, 1 atm — cf. ~0.607 at 25 °C; the 0 °C value is the one used in the lake argument, and the two must not be mixed OBSERVED-REPLICATED (secondary) as above (compilations print 0.6065 with no temperature stated; the T is supplied in-chapter) Measurement at 0 °C, 1 atm outside ~0.55–0.57.
k_snow k_snow ~0.074 (range ~0.05–0.25) W/(m·K) seasonal snow, density-dependent OBSERVED-REPLICATED J Glaciology, 'The thermal conductivity of seasonal snow' Measurement outside range at stated density.
cₚ cₚ 4.181 (4.184 at 20 °C) J/(g·K) liquid water, 25 °C OBSERVED-REPLICATED Wikipedia Table of specific heat capacities (secondary compilation; cites Ashby et al., Young & Geller) Calorimetry outside ±1 %.
cₚ ammonia cₚ ammonia 4.700 — exceeds water J/(g·K) liquid NH₃, 25 °C, saturation pressure ~10 bar (1003 kPa; liquid at 25 °C only under pressure — normal bp −33.34 °C) OBSERVED-REPLICATED (secondary; T and P are not stated in the cited compilation — condition supplied in-chapter. Whether ammonia leads water across the whole liquid range is NOT-VERIFIED) as above Calorimetry at stated T and P showing ammonia < water.
cₚ ethanol / methanol / mercury cₚ ethanol / methanol / mercury 2.440 / 2.140 / 0.1395 J/(g·K) 25 °C OBSERVED-REPLICATED (secondary) as above As above.
ΔH_vap ΔH_vap 2257 (40.65 kJ/mol) kJ/kg 100 °C, normal boiling point OBSERVED-REPLICATED Wikipedia Properties of water; corroborated by steam-table sources in this pass Calorimetry outside ±1 %.
ΔH_vap ethanol ΔH_vap ethanol 38.56 kJ/mol = 837 J/g — near-equal per mole kJ/mol ethanol, normal bp OBSERVED-REPLICATED (tertiary compilation; primary not fetched) surfaced in this pass; falsifier/closure: fetch CRC or NIST Primary value outside ±2 %.
λ_sweat λ_sweat ~2430 J/g the theoretical value at skin temperature — not a measured physiological constant; Havenith measured λ_eff to approach it for evaporation from the skin OBSERVED-REPLICATED (as the value λ_eff approaches at the skin) Havenith et al. 2013, J Appl Physiol 114(6):778–785 A measured λ_eff for skin evaporation outside 2400–2450.
γ γ 71.97 mN/m water–vapour, 25 °C OBSERVED-REPLICATED IAPWS R1-76(2014), recomputed in-chapter from its own correlation → 71.972. The widely-copied 71.99 is Wikipedia Properties of water — a different source Tensiometry outside IAPWS's ±0.5 % below 100 °C.
κ⁻¹ κ⁻¹ 2.71 mm capillary length, √(γ/ρg), 25 °C MODELED Computed in-chapter from γ, ρ(25 °C), g = 9.80665 Arithmetic error.
h_tree,max h_tree,max 112.7 measured; 122–130 predicted m Sequoia sempervirens; ceiling set by leaf water potential, not the meniscus OBSERVED-REPLICATED (measured) / MODELED (ceiling) Koch et al. 2004, Nature 428:851–854 A taller undamaged tree; or a mechanism refuting the water-potential limit.
μ_gas μ_gas 1.8546 D H₂O, gas phase, equilibrium OBSERVED-REPLICATED (secondary compilation; primary not fetched) Wikipedia Properties of water — which states no uncertainty; closure: fetch Clough et al. 1973, J Chem Phys 59:2254, or the NIST/CRC dipole-moment table Primary measurement outside 1.8546 ± 0.001 D.
μ_liquid μ_liquid ~2.6–2.9 (ice Ih ~3.09 ± 0.04) D convention-dependent — no partitioning-free measurement exists OBSERVED-CONTESTED / MODELED reports surfaced in this pass span 2.6–2.9 ± 0.6 A partitioning-independent determination.
ε_r ε_r ≈78.4 dimensionless static dielectric constant, pure water, 25 °C OBSERVED-REPLICATED BNID 115815; formulation: IAPWS R8-97 (Fernández et al., doi 10.1063/1.555997) Value outside ±0.5 at 25 °C.
ΔH_soln(NaCl) ΔH_soln(NaCl) +3.9 — endothermic kJ/mol NaCl in excess water OBSERVED-REPLICATED (secondary/tertiary; primary not fetched) surfaced in this pass; closure: fetch CRC/NIST Calorimetry showing exothermic dissolution.
ΔH_sub(ice Ih) ΔH_sub(ice Ih) 51.059 kJ/mol ice Ih, 0 °C OBSERVED-REPLICATED (tertiary compilation) LSBU Water Structure and Science (Chaplin); primary in Geochim Cosmochim Acta — not fetched in this pass Fetch the primary; value outside ±0.1.
H-bond energy H-bond energy ≤25.5 (bound); ~23.3 (estimate) kJ/mol no standard definition exists OBSERVED-CONTESTED bound computed in-chapter (51.059/2); estimate: Chaplin, 'Water's Hydrogen Bond Strength', arXiv:0706.1355 An agreed operational definition separating the H-bond from dispersion.
τ_HB,osc τ_HB,osc 170 fs underdamped H-bond oscillation period; OH of HOD in D₂O OBSERVED-REPLICATED Fecko et al. 2003, Science 301(5640):1698–1702 Femtosecond IR outside 150–200 fs.
τ_HB,decay τ_HB,decay 1.2 ps decay of vibrational correlations (collective reorganisation) OBSERVED-REPLICATED Fecko et al. 2003 Femtosecond IR outside ~1–1.5 ps.
L_crossover L_crossover ~1 nm hydrophobic small→large regime crossover MODELED (theory + simulation) Huang & Chandler 2000, PNAS 97(15):8324–8327; Lum, Chandler & Weeks 1999, J Phys Chem B 103:4570–4577 A measurement placing the crossover an order of magnitude away.
θ_HOH θ_HOH 104.48 degrees H–O–H bond angle OBSERVED-REPLICATED Wikipedia Properties of water Structural measurement outside ±0.1°.
f_N2 f_N2 78.084 % by volume dry air, sea level OBSERVED-REPLICATED NASA NSSDC Earth Fact Sheet (Colorado mirror) Independent composition measurement outside ±0.01%.
f_O2 f_O2 20.946 % by volume dry air, sea level OBSERVED-REPLICATED as above As above.
f_Ar f_Ar 9,340 (0.934%) ppm by volume dry air OBSERVED-REPLICATED as above As above.
P_O2 P_O2 212 hPa O₂ partial pressure at 1013.25 hPa MODELED Computed in-chapter: 0.20946 × 1013.25 Arithmetic.
[CO₂] [CO₂] 431.44 ppm Mauna Loa monthly mean, June 2026 OBSERVED-REPLICATED NOAA GML Trends in CO₂, updated 05 Jul 2026 Re-read the record; a different published monthly mean for June 2026.
Δ[CO₂]/yr Δ[CO₂]/yr 1.83 ppm/yr June 2025 → June 2026, single-site OBSERVED-REPLICATED NOAA GML, as above As above.
H H 8.43 km RT/(Mg); T=288 K, M=0.028964 kg/mol, g=9.807 MODELED Computed in-chapter Arithmetic; or refute an input.
H (published) H (published) 8.5 km Earth scale height as listed OBSERVED-REPLICATED NASA NSSDC Earth Fact Sheet (mirror) Independent value outside 8–9 km.
M_atm M_atm 5.27 × 10¹⁸ (vs ≈5.15 × 10¹⁸ accepted) kg P₀A/g; gap = mean surface elevation MODELED Computed in-chapter; accepted value not primary-sourced in this pass Fetch a primary M_atm; check the 990 hPa reconciliation.
ρ_air ρ_air 1.204 kg/m³ 20 °C, 101.325 kPa; PM/RT MODELED Computed in-chapter Arithmetic.
μ_air μ_air 1.81 × 10⁻⁵ Pa·s 20 °C, 1 atm OBSERVED-REPLICATED (not primary-sourced in this pass) standard reference; cross-checks against Sutherland's formula → 1.813 × 10⁻⁵ Fetch NIST/CRC; a value outside 1.79–1.84 × 10⁻⁵ refutes.
μ_water μ_water 1.002 × 10⁻³ Pa·s 20 °C, 1 atm OBSERVED-REPLICATED (not primary-sourced in this pass) standard reference (IAPWS-class value) Fetch IAPWS; outside 0.99–1.01 × 10⁻³ refutes.
ρ_water ρ_water 998.2 kg/m³ 20 °C OBSERVED-REPLICATED (not primary-sourced in this pass) standard reference Fetch a primary table.
ν_air ν_air 1.50 × 10⁻⁵ m²/s 20 °C, 1 atm; μ/ρ MODELED Computed in-chapter from the two rows above: 1.81 × 10⁻⁵ / 1.204 = 1.503 × 10⁻⁵ Either input refuted.
ν_water ν_water 1.00 × 10⁻⁶ m²/s 20 °C; μ/ρ MODELED Computed in-chapter; cross-checks vs. published 1.0038 mm²/s at 20.2 °C Either input refuted.
c_air c_air 343.2 m/s 20 °C; √(γRT/M), γ=1.400 MODELED Computed in-chapter Measured c outside 342–344 m/s at 20 °C.
c_water c_water ~1,482 m/s 20 °C OBSERVED-REPLICATED (not primary-sourced in this pass) standard reference Fetch a primary value.
Ω Ω 7.2921 × 10⁻⁵ rad/s Earth sidereal rotation OBSERVED-REPLICATED (not primary-sourced in this pass) standard geodetic constant Fetch IERS conventions.
T_s T_s 288 K mean surface temperature OBSERVED-REPLICATED NASA NSSDC Earth Fact Sheet (mirror) Independent value >2 K off.
T_e T_e 254.6 K [S(1−α)/4σ]^¼; S=1361 W/m², α=0.30 MODELED Computed in-chapter; S and albedo not primary-sourced in this pass — and the NASA sheet cited elsewhere in this chapter disagrees Fetch CERES/TSI primaries; the cited fact sheet's own 0.385 / 1367.6 / 247.3 K disagree with this row and are believed superseded.
greenhouse ΔT greenhouse ΔT 33.4 K T_s − T_e MODELED Computed in-chapter Either input refuted.
O₂ Permian peak O₂ Permian peak ~30 % Phanerozoic maximum, late Palaeozoic MODELED Berner 2006, GCA 70(23):5653–5664; rev. Berner 2009, Am. J. Sci. 309(7):603–606 Recompute the mass balance; a proxy measurement contradicting.
Wingspan M. permiana Wingspan M. permiana ~71 cm Early Permian griffinfly; largest known insect OBSERVED-REPLICATED Guinness World Records (wing impressions, Elmo, Kansas, 1937); widely reported A larger insect wing fossil.
Quetzalcoatlus mass Quetzalcoatlus mass 70 / 200–250 / 544 — a ~8× spread kg 10–11 m wingspan; same fossils, three answers OBSERVED-CONTESTED Chatterjee & Templin 2004 (~70); Paul 2002 / Witton 2008 / Witton & Habib 2010 / Martin & Palmer 2014 (~200–250, the consensus cluster); Henderson 2010 (~544) — via secondary reports A method reconciling the three; do not quote one as 'the' mass — and do not round the ends inward, which shrinks the finding.
Pelagornis wingspan Pelagornis wingspan 6.06–7.38 m depends on feather-reconstruction method OBSERVED-REPLICATED Ksepka 2014, PNAS, doi:10.1073/pnas.1320297111 Re-measure; a span outside the range.
Pelagornis mass Pelagornis mass ~22–40 kg regression-dependent; exceeds the flapping ceiling — it soared OBSERVED-REPLICATED Ksepka 2014 As above.
T_c,☉ T_c,☉ 1.54 × 10⁷ K standard solar model, current epoch MODELED arXiv:2501.09971 Table 1; cf. Salmon et al. 2021, A&A 651, A106 An SSM variant outside 1.4–1.7 × 10⁷ K reproducing helioseismology + neutrinos.
ρ_c,☉ ρ_c,☉ 149 g/cm³ standard solar model, current epoch MODELED arXiv:2501.09971 Table 1 As above.
P_c,☉ (SSM) P_c,☉ (SSM) 2.3 × 10¹⁶ Pa standard solar model, current epoch MODELED arXiv:2501.09971 Table 1 An SSM variant outside this range reproducing helioseismology + neutrinos.
t_☉,formation t_☉,formation 4.567 × 10⁹ yr Pb–Pb dating of CAIs OBSERVED-REPLICATED Connelly et al. 2012, Science 338:651–655 Independent radiometric dating >1% off.
Φ(CNO) final Φ(CNO) final 6.7 (+1.2 / −0.8) × 10⁸ cm⁻² s⁻¹ solar CNO neutrinos; CID over the complete 2007–2021 dataset + an improved Phase-III spectral fit OBSERVED-REPLICATED Borexino Collaboration 2023, Phys Rev D 108:102005, arXiv:2307.14636 An independent detector outside the interval.
t½(⁸Be) t½(⁸Be) ~10⁻¹⁶ s ⁸Be ground state, unbound OBSERVED-REPLICATED (standard nuclear data; not primary-sourced in this pass) standard nuclear data tables Measured lifetime >10× off.
E_x(Hoyle) modern E_x(Hoyle) modern 7,654.07 ± 0.19 keV ¹²C second excited state, 0⁺ OBSERVED-REPLICATED NNDC, via Freer & Fynbo 2014, Prog Part Nucl Phys 78:1–23 A measurement outside ±0.19 keV.
BE/A(⁶²Ni) BE/A(⁶²Ni) 8.7945 MeV/nucleon highest known binding energy per nucleon OBSERVED-REPLICATED standard nuclear mass tables A nuclide measured higher.
BE/A(⁵⁸Fe) BE/A(⁵⁸Fe) 8.7922 MeV/nucleon second OBSERVED-REPLICATED as above As above.
BE/A(⁵⁶Fe) BE/A(⁵⁶Fe) 8.7903 MeV/nucleon third — but lowest mass per nucleon (a different quantity) OBSERVED-REPLICATED as above Show ⁵⁶Fe has the highest BE/A, or that mass/nucleon and BE/A are the same quantity.
ΔBE/A (⁶²Ni − ⁵⁶Fe) ΔBE/A (⁶²Ni − ⁵⁶Fe) 4.2 (0.048%) keV/nucleon the whole 'iron peak' spread MODELED Computed in-chapter from the rows above Arithmetic error.
t½(⁵⁶Ni → ⁵⁶Co) t½(⁵⁶Ni → ⁵⁶Co) ~6 d powers the early Type Ia light curve OBSERVED-REPLICATED (standard nuclear data; not primary-sourced in this pass) standard nuclear data tables Measured half-life >10% off.
t½(⁵⁶Co → ⁵⁶Fe) t½(⁵⁶Co → ⁵⁶Fe) ~77 d powers the Type Ia light-curve tail OBSERVED-REPLICATED (standard nuclear data; not primary-sourced in this pass) standard nuclear data tables As above.
L_Edd L_Edd 1.26 × 10³¹ (M/M☉) ≈ 3.3 × 10⁴ (M/M☉) L☉ W 4πGMm_p c/σ_T; spherical, steady, ionised H, Thomson opacity only MODELED Eddington 1926, The Internal Constitution of the Stars; arithmetic computed in-chapter Arithmetic error; or a steady spherical source persistently above it.
Y_p Y_p ~0.245 mass fraction primordial helium, BBN/CMB concordance OBSERVED-REPLICATED (standard reference; not primary-sourced in this pass) BBN/CMB literature An independent determination outside ~0.24–0.25.
Δc_s/c_s Δc_s/c_s 0.10 % (rms fractional) SSM vs helioseismic inversion, 0.05–0.95 R☉ OBSERVED-REPLICATED (inversion) / MODELED (the model side) Bahcall, Pinsonneault & Basu 2001, ApJ 555:990 An independent inversion disagreeing at >0.5%.
Ṗ_b,obs/Ṗ_b,GR Ṗ_b,obs/Ṗ_b,GR 0.9983 ± 0.0016 dimensionless PSR B1913+16; 9,257 TOAs over 35 yr OBSERVED-REPLICATED Weisberg & Huang 2016, ApJ 829:55, doi:10.3847/0004-637X/829/1/55 A ratio outside the interval on longer baselines.
M_psr / M_comp M_psr / M_comp 1.438 ± 0.001 / 1.390 ± 0.001 M☉ PSR B1913+16 OBSERVED-REPLICATED Weisberg & Huang 2016 As above.
Au in AT2017gfo Au in AT2017gfo NOT DETECTED; M_Au ≲ 10⁻², M_Pt ≲ few × 10⁻³ M☉ (upper limits) 'no platinum or gold signatures are prominent in the ejecta' NOT-MEASURED (upper limits only) Gillanders et al. 2021, MNRAS 506(3):3560–3577, doi:10.1093/mnras/stab1861 A spectroscopic identification of Au or Pt in a kilonova.
rise rise ~3.4 (0.34) Å (nm) /bp B-DNA OBSERVED-REPLICATED Potaman & Sinden, NCBI Bookshelf NBK6545; Watson & Crick 1953, Nature 171:737–738 Structural measurement outside 3.3–3.5 Å under B-form conditions.
bp/turn (fibre) bp/turn (fibre) 10.0 bp B-form fibre diffraction OBSERVED-REPLICATED Watson & Crick 1953; as compared in Wang 1979 Re-analysis of fibre data giving ≠10.
bp/turn (solution) bp/turn (solution) 10.4 ± 0.1 bp B-DNA free in solution, physiological; topoisomer gel OBSERVED-REPLICATED Wang 1979, PNAS 76(1):200–203, doi:10.1073/pnas.76.1.200 Repeat topoisomer ladder; value outside 10.3–10.5.
bp/turn (Rhodes & Klug) bp/turn (Rhodes & Klug) 10.6 ± 0.1 bases DNase I cutting periodicity on DNA immobilised on three surfaces; the authors identify the value with the repeat in solution OBSERVED-REPLICATED Rhodes & Klug 1980, Nature 286(5773):573–578, PMID 7402337 Repeat digestion periodicity outside 10.5–10.7.
bp/turn '10.5' bp/turn '10.5' 10.5 bp quoted everywhere; printed by Potaman & Sinden Table 1 as B-DNA residues/turn; sits between two unreconciled primaries NOT-SOURCED in this pass (value printed in source; provenance not traced) Potaman & Sinden, NBK6545 Table 1 Establish whether 10.5 is an independent determination, a rounding, or a midpoint of 10.4 and 10.6.
diameter diameter 23 (A) / ~20 (B) / 18 (Z) Å A-, B-, Z-DNA OBSERVED-REPLICATED Potaman & Sinden, NBK6545 Table 1 Structural measurement outside range.
groove widths (12/6) groove widths (12/6) major ~12, minor ~6 Å B-DNA, convention not sourced here NOT-SOURCED in this pass not confirmed here — NBK6545 Table 1 has no groove-width row; the prior attribution to it was fabricated (recorded in-chapter) Fetch a primary printing these values and naming the measurement convention.
groove widths (22/12) groove widths (22/12) 22 / 12 Å widely repeated textbook pair — different convention NOT-SOURCED in this pass not confirmed here Fetch a primary source and name the measurement convention.
ℓ_p ℓ_p ~50 (~150) nm (bp) dsDNA, ~0.1 M NaCl, ~20–25 °C; tweezers + AFM consensus OBSERVED-REPLICATED Peters & Maher 2010, Q Rev Biophys 43(1):23–63, PMID 20478077 Independent single-molecule measurement outside 45–55 nm at the stated temperature and ionic strength.
ℓ_p vs temperature ℓ_p vs temperature 53.2 nm (5 °C) → 42.5 nm (42 °C) nm TBE + 10 mM MgCl₂ — a different ionic condition from the row above; j-factor + linking-number methods. A condition on the 50 nm consensus, not support for it: at ≥37 °C it sits at or below that row's own falsifier band OBSERVED-REPLICATED Geggier, Kotlyar & Vologodskii 2011, NAR 39(4):1419–1426, PMID 20952402 A measurement showing ℓ_p temperature-independent across 5–42 °C.
I/bp I/bp 2 bits/bp log₂(4); upper bound, assumes no correlation MODELED Computed in-chapter Arithmetic error (the bound itself is definitional).
G_human G_human 3,054,815,472 (+16,569 mtDNA) bp T2T-CHM13, gapless, chr1–22 + X OBSERVED-REPLICATED Nurk et al. 2022, Science 376:44–53, doi:10.1126/science.abj6987 Independent T2T assembly differing >0.1%.
f_coding f_coding 1–2 (exome ~1.5) % of genome human OBSERVED-REPLICATED Piovesan et al. 2019, BMC Res Notes, doi:10.1186/s13104-019-4343-8 Annotation revision moving coding fraction outside 1–2%.
pg→bp pg→bp 1 pg = 0.978e9 bp flow-cytometry conversion OBSERVED-REPLICATED Doležel et al. 2003, Cytometry A, doi:10.1002/cyto.a.10013 Re-derivation of nucleotide-pair molecular weight.
G_onion G_onion 16.75 pg ≈ 16.4 (≥95% repetitive) Gbp Allium cepa 1C OBSERVED-REPLICATED onion assembly literature (PMC8496297; PMC11865573) + Doležel conversion Flow-cytometry re-measurement outside 15–18 Gbp.
G_lungfish G_lungfish 91 (~90% repeat) Gbp Lepidosiren paradoxa — largest sequenced animal genome OBSERVED-REPLICATED Schartl et al. 2024, Nature 634:96–103, doi:10.1038/s41586-024-07830-1 Independent assembly differing >10%.
G_fern G_fern 160.45 Gbp/1C Tmesipteris oblanceolata — current eukaryotic record OBSERVED-REPLICATED Fernández et al. 2024, iScience, doi:10.1016/j.isci.2024.109889 Independent flow cytometry differing >10%.
G_Paris G_Paris 152.23 pg ≈ 149 Gbp Paris japonica 1C OBSERVED-REPLICATED Pellicer et al. 2010, Bot J Linn Soc 164(1):10, doi:10.1111/j.1095-8339.2010.01072.x Re-measurement outside range.
nucleosome nucleosome 146 bp @ 2.8 Å; 147 bp @ 1.9 Å; 1.65 superhelical turns; histone octamer bp crystal structures — different crystals, both real OBSERVED-REPLICATED Luger et al. 1997, Nature 389:251–260, doi:10.1038/38444; Richmond & Davey 2003, Nature 423:145, doi:10.1038/nature01595 A re-refinement changing the wrap length.
μ_germline μ_germline 1.20e-8 per nt per generation (mean paternal age 29.7; +~2 mutations/yr; paternal doubling ~16.5 yr) human trios — different denominator from ε OBSERVED-REPLICATED Kong et al. 2012, Nature 488(7412):471–475, doi:10.1038/nature11396 Independent trio study outside range at matched paternal age.
v_fork,ec v_fork,ec ~600 (classic ~1,000) bp/s E. coli, in vivo average OBSERVED-REPLICATED BNID 109251; Milo & Phillips In vivo measurement outside 400–1,000 bp/s.
v_fork,euk v_fork,euk 4–40 (~1 kb/min) bp/s eukaryotic replisome OBSERVED-REPLICATED BNID 104930, 104935, 104936, 104937 Outside range.
T_S T_S ~10 hours human S phase OBSERVED-REPLICATED BNID 103742, 103741, 102204 Cell type outside range.
N_origins (measured) N_origins (measured) 1,000–100,000 (Drosophila ~10,000) origins human OBSERVED-REPLICATED BNID 107654, 109283 A measurement excluding the predicted band.
u u 62 (low visc) / 65 (high visc) µm/s migrating human sperm, 37 °C, n = 16 / 19 OBSERVED-REPLICATED Smith et al. 2009, Cell Motil Cytoskeleton 66(4):220–236, DOI 10.1002/cm.20345 Repeat high-frame-rate imaging of the migrating cohort; a mean outside 40–90 µm/s refutes.
λ λ 39 (low visc) / 18 (high visc) µm human sperm flagellar wavelength, 37 °C OBSERVED-REPLICATED Smith et al. 2009 Outside range at stated viscosity.
c_wave c_wave 890 (low visc) / 200 (high visc) µm/s wavespeed = f × λ OBSERVED-REPLICATED Smith et al. 2009 Recompute; c ≠ fλ under stated conditions.
L_flag L_flag ~41 µm human flagellum, tethered-cell imaging OBSERVED-REPLICATED Saggiorato et al. 2017 Independent measurement >1.5× off.
µ_buffer µ_buffer 0.7 (0.73 ± 0.01 for HTF) mPa·s aqueous buffer, 37 °C OBSERVED-REPLICATED Saggiorato et al. 2017 Rheometry outside range at 37 °C.
µ_mucus µ_mucus ~0.14 (analogue) / ~0.2 (midcycle, Day 0) / ~0.68 (Day 5) Pa·s Maxwell fit to measured G′/G″ at ~5 Hz, 37 °C MODELED Smith et al. 2009 (rheometry + fit); mucus moduli from Wolf et al. 1977, Fertil Steril 28:47–52 A direct steady-shear viscosity of periovulatory mucus outside 0.1–1 Pa·s.
d_coast d_coast ~0.45 Å (4.5 × 10⁻¹¹ m), τ ≈ 0.7 µs m Stokes coasting, τ=m/(6πµa); V≈17 µm³, ρ_cell≈1.1×10³, a≈2 µm MODELED Computed in-chapter (inputs order-of-magnitude) Recompute; a measured coasting distance >1 nm refutes.
n_dynein n_dynein ~67,500 total; ~15,000 active/beat motors sea-urchin sperm flagellum; active count is a hypothesis (67,500 × 2/9) OBSERVED-REPLICATED (total, cryo-ET) / HYPOTHESIZED (active fraction) Chen et al. 2015, Biophys J 109:2562–2573 Count in situ; an active fraction ≠ ~2/9.
d_dynein d_dynein ~8 nm/power stroke axonemal dynein, single-molecule OBSERVED-REPLICATED Chen et al. 2015 and refs therein Different step periodicity.
n_ATP/beat n_ATP/beat (2.3 ± 0.2) × 10⁵ (low visc) → (3.2 ± 0.5) × 10⁵ (0.5 % MC) ATP/beat demembranated sea-urchin sperm axoneme, single-cell, [ATP] = 20 µM, viscosity as stated — this quantity is not viscosity-independent OBSERVED-SINGLE (single study) Chen et al. 2015 Bulk S. purpuratus gives ~1 × 10⁵/beat — already a 2.3× discrepancy; carry it.
r_ATP r_ATP (2.4 ± 0.3) × 10⁶ active; (9.2 ± 0.2) × 10⁵ inactive ATP/s demembranated Lytechinus sperm axoneme OBSERVED-SINGLE (single study) Chen et al. 2015 Independent single-cell measurement >2× off.
ε_hydro ε_hydro 0.004 (low visc) → 0.013 (high visc) dimensionless demembranated sea-urchin axoneme, 20 µM ATP, buffer vs 0.5 % MC OBSERVED-SINGLE (single study — not replicated) Chen et al. 2015, Table S1 Independent replication >3× off; not transferable to intact human sperm.
ε_chemo ε_chemo 0.34 → 0.6 dimensionless as above OBSERVED-SINGLE (single study — not replicated) Chen et al. 2015 As above.
ε_swim ε_swim 0.001 → 0.008 dimensionless as above; ε_swim = ε_chemo · ε_hydro OBSERVED-SINGLE (single study — not replicated) Chen et al. 2015 As above.
n_mito n_mito ~50–75 (~1 mtDNA each) per cell human sperm midpiece OBSERVED-SINGLE (via review; primary count not retrieved in this pass) Hirata et al. 2002, Reprod Med Biol 1(2):41–47 Direct EM/qPCR count outside range.
toroid toroid ~50 (up to ~60) kb DNA per toroid protamine–DNA toroid OBSERVED-REPLICATED Hud et al. 1995, as reviewed in Balhorn 2007; single-molecule receipt for the toroid mechanism (λ-phage DNA in an optical trap — not sperm chromatin, not a compaction ratio): Brewer, Corzett & Balhorn 1999, Science 286(5437):120–123 Structural measurement outside range.
min. gradient min. gradient 0.8 fM/µm A. punctulata, resact OBSERVED-SINGLE (single group — not independently replicated) Kashikar et al. 2012, J Cell Biol 198(6):1075–1091 Independent measurement >10× off.
N_ejac N_ejac median ~255 × 10⁶; 5th-centile 39 × 10⁶ sperm/ejaculate WHO reference population (TTP ≤ 12 months) OBSERVED-REPLICATED Cooper et al. 2010, Hum Reprod Update 16(3):231–245; WHO 2021, 6th ed. Re-derive from the reference cohort.
N_tube N_tube median 251 (range 79–1,386) sperm in both Fallopian tubes 10 parous women, ~18 h post-insemination, tubes ligated + flushed OBSERVED-SINGLE (single study, n = 10 — not replicated) Williams et al. 1993, Hum Reprod 8(12):2019–2026 An independent flush study giving a median outside ~50–2,000.
T_colony T_colony ~25 years Pogonomyrmex barbatus, single-queen founding OBSERVED-REPLICATED Gordon et al. 2011, Behav Ecol 22(2):429–435, quoting Gordon 1991 Long-term census showing mean colony persistence outside ~15–30 yr.
T_worker,ext T_worker,ext ≤ 33 (the paper's summary sentence, covering both spp.: 'about 30') days after marking P. barbatus exterior workers, field, 3,521 marked across 38 mature colonies, Rodeo NM, Jul–Aug 1987, checked once daily; max 33 d = a nest-maintenance worker OBSERVED-REPLICATED Gordon & Hölldobler 1987, Psyche 94:341–346 Authors name two biases (paint wear-off; marked ants alive inside) that would underestimate — a mark–recapture design controlling both, finding >>33 d, moves this row.
t½_forager t½_forager 4.2 days (half-life, marked foragers) Cataglyphis bicolor, Southern Tunisia; constant 16.4%/day loss; single study, not independently replicated OBSERVED-SINGLE — primary read directly in this pass. The widely-cited secondary is wrong: Gordon & Hölldobler 1987 print a '6 day half-life', which is the mean life expectancy wearing a half-life label Schmid-Hempel & Schmid-Hempel 1984, Ins Soc 31:345–360, DOI 10.1007/BF02223652 An independent marked-forager census in C. bicolor returning a half-life >2× off.
E[life]_forager E[life]_forager 6.1 days (mean life expectancy) C. bicolor, Southern Tunisia, same study and same 16.4%/day loss; single study, not independently replicated OBSERVED-SINGLE — primary read directly. This is the number the secondary literature misprints as a 6-day 'half-life' Schmid-Hempel & Schmid-Hempel 1984 As above.
N_colony N_colony 10,000–12,000 (young: 2,000–10,000) workers P. barbatus at reproductive age (5 yr) OBSERVED-REPLICATED Gordon et al. 2011 Census outside range for mature colonies.
N_brain (IF) N_brain (IF) 7.02 × 10⁴ ± 2.4 × 10⁴ (s) brain nuclei Novomessor spp. (desert ant), isotropic fractionator — the method of the 32 spp. / 7 superfamilies survey. The SD is ~⅓ of the mean and travels with the value OBSERVED-REPLICATED Godfrey, Swartzlander & Gronenberg 2021, Proc R Soc B 288:20210199 Independent count >2× off under the same method. Not a Pogonomyrmex number — do not transfer across genera.
N_brain (sectioned) N_brain (sectioned) ≈ 9 × 10⁴ brain nuclei Novomessor spp., sectioned brains — the paper's method-validation cross-check on the IF adaptation, not an IF count; agrees with the IF value within ~1 SD OBSERVED-REPLICATED Godfrey et al. 2021 As above. Never print this figure under the IF label.
release rate release rate 0.25 ± 0.10 pg·cm⁻¹·s⁻¹ (Z)-9-hexadecenal from filter-paper trails, I. humilis OBSERVED-REPLICATED Van Vorhis Key & Baker 1982 Independent measurement >2× off.
t_decay,attract t_decay,attract 33 min (95% CI of fitted curve reaching no-effect) Monomorium pharaonis, ECF paper, 10 colonies (~1,500 workers) OBSERVED-REPLICATED Robinson et al. 2008, Insectes Sociaux 55:246–251 Replication on the same substrate outside CI.
t_decay,repel t_decay,repel 78 min (same criterion) M. pharaonis, repellent 'no entry' pheromone, 11 qualifying trials OBSERVED-REPLICATED Robinson et al. 2008 As above.
colony mass range colony mass range 0.0017 → 3,850 g Solenopsis morphospecies → Macrotermes bellicosus OBSERVED-REPLICATED Hou et al. 2010
M_adult,Trex,published M_adult,Trex,published 6,000–8,000; 'Sue' perhaps ~9,500 kg adult T. rex — the authors' own stated conclusion OBSERVED-CONTESTED Hutchinson et al. 2011, PLoS ONE 6(10):e26037, abstract An independent volumetric study concluding outside this band.
M(PR2081)_min M(PR2081)_min 9,502 kg T. rex 'Sue', volumetric minimal model — authors' preferred region OBSERVED-CONTESTED Hutchinson et al. 2011, Table 6 Independent volumetric reconstruction outside ~8,000–11,000 kg.
M(PR2081)_max M(PR2081)_max 18,489 kg T. rex 'Sue', volumetric maximal model — authors judge 'less plausible' OBSERVED-CONTESTED Hutchinson et al. 2011, Table 6 As above.
ASP ASP ~50–60%, up to 79% (Sauroposeidon) % vertebral volume as air adult neosauropod cervicals OBSERVED-REPLICATED Wedel 2005, as reported by Schwarz-Wings et al. 2009, Proc R Soc B 277(1678):11–17 (they cite it to Wedel; they measured no Sauroposeidon and generated no ASP data — do not name the quoter as the source) Re-measure Wedel's CT dataset; ASP outside range.
ASP (2nd) ASP (2nd) 0.50–0.70; specific gravity to 0.2 (vs compact bone 1.8–2.0) dimensionless sauropod cervicals OBSERVED-REPLICATED Taylor & Wedel 2013, PeerJ 1:e36, DOI 10.7717/peerj.36 As above.
T_body,sauropod T_body,sauropod 36–38 °C large Jurassic sauropod teeth, clumped-isotope (¹³C–¹⁸O) OBSERVED-REPLICATED Eagle et al. 2011, Science 333(6041):443–445, DOI 10.1126/science.1206196 Independent thermometry outside range.
L_neck,max L_neck,max ~15.1 m Mamenchisaurus sinocanadorum; >6× giraffe OBSERVED-CONTESTED Moore et al. 2023, J Syst Palaeontol 21(1), DOI 10.1080/14772019.2023.2171818 New cervical material; note 2013 estimate was ~12 m.
L_neck,giraffe L_neck,giraffe 2.4 m world-record bull giraffe OBSERVED-REPLICATED Taylor & Wedel 2013 A longer measured giraffe neck.
n_cervical n_cervical sauropods 13–17 (19 in M. hochuanensis); mammals exactly 7 (sloths/sirenians excepted) count OBSERVED-REPLICATED Taylor & Wedel 2013 A mammal outside the exceptions with ≠7.
dP/dz dP/dz 77.3 (78.0 at ρ=1060) mmHg per metre ρg, ρ_blood ≈ 1050 kg·m⁻³, g = 9.81 — a definition (hydrostatics), not a measurement MODELED Computed in-chapter. The ~77 mmHg/m quoted in the giraffe literature is this same ρg, not an independent measurement of it — no cross-check is claimed Arithmetic error, or ρ_blood refuted.
MAP_sauropod MAP_sauropod 750 (= 700 static column + ~50 perfusion) mmHg ~9 m raised head MODELED Seymour 2009, Biol Lett 5(3):317–319 — the in-chapter arithmetic REPRODUCES his construction using his own +50 term: a consistency check on transcription, NOT an independent landing Re-derive; a different ρ or geometry.
MAP_giraffe,heart MAP_giraffe,heart 185 ± 41.6 mmHg giraffe, at heart level OBSERVED-REPLICATED Mitchell et al. 2006, J Exp Biol 209(13):2515 Independent catheterisation outside range.
MAP_giraffe,head MAP_giraffe,head 100.3 ± 20.9 mmHg giraffe, at head OBSERVED-REPLICATED Mitchell & Skinner 1993, via Mitchell et al. 2006 As above.
M_blue,measured M_blue,measured t mass of the largest blue whale NOT-MEASURED Motani & Pyenson 2024, PeerJ 12:e16978 Weigh one intact.
M_blue,piece M_blue,piece ≥136.4 t 27.1 m female, weighed in parts (fluid lost) OBSERVED-SINGLE Winston 1950, via Motani & Pyenson 2024 Re-weigh under controlled loss accounting.
M_blue,regr M_blue,regr 234 (95% CI 187–294); fluid-corrected 252 (CI 201–306) @ 7% blood loss, 272 (CI 217–342) @ 14% t 33 m blue whale, length→mass regression. The two fluid-corrected values are separate estimates under separate blood-loss assumptions — never merge them into one interval; 'CI 201–342' is a hull of two intervals, not an interval MODELED Motani & Pyenson 2024 New regression on a larger measured sample; an independent fluid-loss accounting outside 7–14%.
M_blue,vol M_blue,vol 266–279 t 3D volumetric model MODELED Motani & Pyenson 2024 Independent volumetric model outside range.
L_blue,max L_blue,max 33.26 m longest reliably measured blue whale OBSERVED-SINGLE Risting 1928, via Motani & Pyenson 2024 A longer verifiable measurement.
TE_trophic TE_trophic 10 % per trophic level 48 trophic models, 6 aquatic ecosystem types; re-estimated, not assumed OBSERVED-REPLICATED Pauly & Christensen 1995, Nature 374:255–257 Re-estimate from independent models outside ~5–15%.
TL_range TL_range 1.0 (edible algae) – 4.2 (tunas) fractional trophic level 39 commodity groups, global catch 94.3 Mt/yr 1988–91 OBSERVED-REPLICATED Pauly & Christensen 1995 Re-assign trophic levels.
[Mb]_cetacean [Mb]_cetacean 1.81–5.78 g Mb / 100 g wet muscle cetaceans, body mass 70–80,000 kg OBSERVED-REPLICATED Noren & Williams 2000, Comp Biochem Physiol A 126(2):181–191 Measurement outside range under stated method.
[Mb]_dolphin,max [Mb]_dolphin,max ~6.3 g Mb / 100 g striped dolphin, epaxial middle OBSERVED-SINGLE Arregui et al. 2021, Animals 11(2):451 Independent assay >2× off.
[Mb]_terrestrial [Mb]_terrestrial <~0.5 (ratio 10–30× divers:non-divers) g Mb / 100 g non-diving mammals — Kooyman 1989 primary not read here OBSERVED-REPLICATED (secondary attribution) Kooyman 1989, cited in the comparative literature Fetch Kooyman 1989; measure human muscle Mb.
[Mb]_human [Mb]_human g Mb / 100 g human skeletal muscle NOT-MEASURED not sourced in this pass Fetch a primary value.
d_collapse,N₂ d_collapse,N₂ ~70 m Tursiops truncatus, live, inferred from N₂ washout OBSERVED-SINGLE Ridgway & Howard 1979, Science 206(4423):1182–1183 Repeat washout; a different inferred depth.
d_collapse,CT d_collapse,CT range 58 (grey seal, 50% TLC) – 133 (harbour porpoise, 100% TLC); per-specimen common-dolphin values NOT-CONFIRMED m post-mortem hyperbaric CT, extrapolated to zero gas volume — vessel rated only to 170 m depth-equivalent, so these are extrapolations, not readings. Every value carries a TLC condition. Do not average with the row above OBSERVED-CONTESTED Moore et al. 2011, J Exp Biol 214:2390–2397 (abstract only — Table 2 not read in this pass) In-vivo imaging under pressure reconciling both; fetch Table 2 for per-specimen values.
d_dive,max d_dive,max 2,992 m Ziphius cavirostris, mammalian depth record OBSERVED-SINGLE Schorr et al. 2014, PLOS ONE 9(3):e92633 A deeper tagged mammalian dive.
ΔdB_air/water ΔdB_air/water 61.58 dB 10·log₁₀(20² × 3600): reference-pressure ratio × impedance ratio MODELED (arithmetic) Computed in-chapter; cross-checked against Møhl et al. 2003's own 235→173 conversion Arithmetic error; a different impedance ratio.
c_sound c_sound 1477 (surface) → 1468 (500 m) m/s Norwegian coastal water, measured profile OBSERVED-SINGLE Møhl et al. 2003 Independent CTD profile.
z_SOFAR z_SOFAR ~750–1,200 (midlat); near-surface polar m deep sound channel axis OBSERVED-REPLICATED (secondary source in this pass) Ewing & Worzel 1948, GSA Memoir 27; depth via secondary Fetch a primary sound-speed climatology.
r_SOFAR,demo r_SOFAR,demo up to 900 nmi (~1,700 km) km 1944 R/V Saluda explosive-charge demonstration OBSERVED-SINGLE Ewing & Worzel 1948 Read the primary; a different demonstrated range.
c c ≈343 m/s air, 20 °C, 1 atm; c ≈ 331.3·sqrt(1+T/273.15) OBSERVED-REPLICATED (standard reference; not primary-sourced in this pass) standard acoustics Fetch a primary reference; a measured c outside 330–350 at stated conditions.
alpha_atm (25→50 kHz) alpha_atm (25→50 kHz) 0.7 → 1.7 dB/m 25 → 50 kHz, 20 °C, 50% RH OBSERVED-REPLICATED Jakobsen, Brinkløv & Surlykke 2013, Front Physiol 4:89, from Lawrence & Simmons 1982; ANSI 1995 Re-measure at stated T/RH; values >2× off.
alpha_atm (45→90 kHz) alpha_atm (45→90 kHz) 1.4 → 4 dB/m 45 → 90 kHz, 25 °C, 80% RH OBSERVED-REPLICATED as above As above.
A_MEM A_MEM 17–25 dB attenuation of self-generated signal middle-ear muscle contraction during emission OBSERVED-REPLICATED (spread real; some sources report ~20–30) Suga & Jen 1975; Henson 1965 (cited by, not read here) Measure with stapedius intact vs ablated.
AGC AGC ~6 dB sensitivity drop per halving of target distance approach phase, attributed to middle-ear muscles OBSERVED-REPLICATED Jakobsen et al. 2013, citing Suga & Jen 1975 Measure receiver gain vs range and find no schedule.
p@140dB p@140dB 200 Pa (≈0.2% of 1 atm) p = 20e-6 × 10^(SL/20) at 0.1 m MODELED Computed in-chapter Arithmetic error.
miss_distance miss_distance ~15–20 cm jammed bats; matches ranging-interference prediction OBSERVED-REPLICATED Corcoran, Barber, Hristov & Conner 2011, J Exp Biol 214:2416–25 A method discriminating the three hypotheses and favouring phantom echo or masking.
d_detect,moth d_detect,moth 20–25 (E. fuscus) vs <1 (E. maculatum) m average noctuoid moth's detection distance OBSERVED-REPLICATED Fullard & Dawson 1997 Neurophysiology giving a different threshold; a moth detecting E. maculatum at >5 m.
N_cells,Bianconi N_cells,Bianconi 3.72 × 10¹³ cells standard adult; organ-by-organ bibliographic sum MODELED Bianconi et al. 2013, Ann Hum Biol 40(6):463–471 Independent systematic count outside 2–6 × 10¹³.
N_cells,Sender N_cells,Sender 3.0 × 10¹³ (2% est. unc.; 14% CV) cells 70 kg reference male MODELED Sender, Fuchs & Milo 2016, PLoS Biol 14(8):e1002533 Recount outside stated uncertainty.
N_bacteria N_bacteria 3.8 × 10¹³ (25% est. unc.; 52% SD) cells 70 kg male, mostly colon MODELED Sender et al. 2016 Independent estimate outside stated band.
R_B/H R_B/H 1.3 (25% unc.; 53% variation) dimensionless all human cells as denominator MODELED Sender et al. 2016 Recount outside band.
R_B/H,nucleated R_B/H,nucleated ~8:1 dimensionless nucleated human cells as denominator MODELED Computed in-chapter from Sender et al. 2016 Arithmetic error.
m_microbiota m_microbiota ~0.2 (0.3% of body mass) kg wet weight 70 kg male MODELED Sender et al. 2016 Direct measurement >2× off.
V̇O₂,basal V̇O₂,basal 250 mL O₂/min 70 kg man, basal state OBSERVED-REPLICATED Clarke & Sokoloff, Basic Neurochem, 6th ed. 1999 Indirect calorimetry >20% off.
k_O₂ k_O₂ 4.82 (≈20.1 kJ/L) kcal/L O₂ mixed diet, RQ ≈ 0.82 (fat 4.68, CHO 5.03) OBSERVED-REPLICATED standard indirect calorimetry Bomb-calorimetry value outside 4.6–5.1.
P_BMR P_BMR ~84 (≈1,735 kcal/day) W 70 kg man; from V̇O₂ × k_O₂ MODELED Computed in-chapter from Clarke & Sokoloff Arithmetic error, or V̇O₂ refuted.
p_body p_body ~1.2 W/kg 84 W / 70 kg MODELED Computed in-chapter Arithmetic error.
N_neurons N_neurons 86.1 ± 8.1 × 10⁹ neurons adult male human brain, isotropic fractionator OBSERVED-REPLICATED Azevedo et al. 2009, J Comp Neurol 513:532–541 Independent count outside ±2 SD.
N_nonneuronal N_nonneuronal 84.6 ± 9.8 × 10⁹ cells same OBSERVED-REPLICATED Azevedo et al. 2009 As above.
R_glia:neuron R_glia:neuron ~1:1 dimensionless whole human brain OBSERVED-REPLICATED Azevedo et al. 2009 A count returning ≥3:1 by an unbiased method.
f_cortex f_cortex 19 % of all brain neurons adult male human, isotropic fractionator — the human-specific unbiased count OBSERVED-REPLICATED Azevedo et al. 2009 (abstract: 'With only 19% of all neurons located in the cerebral cortex') Unbiased count outside 17–21%.
N_synapses N_synapses 1.64 × 10¹⁴ (164 × 10¹², CV = 0.17; textbook summaries span 0.6–2.4 × 10¹⁴) synapses human neocortex, n = 5 young male autopsy brains, unbiased stereology OBSERVED-SINGLE (single study, n = 5; no independent replication located) Tang et al. 2001, Synapse 41(3):258–273 — primary abstract read this pass An independent stereological count outside 0.6–2.4 × 10¹⁴; any replication at n > 5.
f_brain,mass f_brain,mass ~2 % of body mass ~1.4–1.5 kg / 70 kg OBSERVED-REPLICATED standard anatomy; Herculano-Houzel 2012 Measurement outside 1.5–2.5%.
f_brain,O₂ f_brain,O₂ ~20 % of resting O₂ consumption 70 kg man, basal OBSERVED-REPLICATED Clarke & Sokoloff 1999 CMRO₂ measurement outside 15–25%.
P_brain,CMRO₂ P_brain,CMRO₂ ~17 (≈347 kcal/day) W from 20% × 250 mL O₂/min MODELED Computed in-chapter from Clarke & Sokoloff Arithmetic error.
P_brain,perneuron P_brain,perneuron ~25 (516 kcal/day; 25% of TDEE) W 86 × 10⁹ × 6 kcal/d per 10⁹ neurons MODELED Herculano-Houzel 2012 Direct human measurement settling the 1.5× gap.
N_beats,mammal N_beats,mammal 7.3 ± 5.6 × 10⁸ (CV ≈ 77%) beats/lifetime 15 mammal species, excluding humans. Provenance, per Levine's own Fig. 1 caption — 'Most coordinates represent average values (4–6)': refs 4–6 are Spector, Handbook of Biological Data (1956); Encyclopaedia Britannica (1971); and White et al. (1938). Species lifespan/HR coordinates not independently traced OBSERVED-CONTESTED (a loose 'invariant', on averaged and untraced coordinates) Levine 1997, J Am Coll Cardiol 30:1104–1106 Expand the species set; a tighter or looser CV; trace the coordinates to primary measurements.
N_beats,human N_beats,human ~2.9 × 10⁹ (2.945 × 10⁹; ~4× mean, ~4 SD out) beats/lifetime 70 bpm × 80 yr MODELED Levine 1997 states ~3 × 10⁹ himself at these inputs — recomputed in-chapter as 2.945 × 10⁹ Arithmetic error; Levine's own stated figure moving.
E_threshold E_threshold 2.1–5.7 × 10⁻¹⁰ ergs = 54–148 quanta at the cornea blue-green, optimal conditions OBSERVED-REPLICATED Hecht, Shlaer & Pirenne 1942, J Gen Physiol 25(6):819–840 Replication outside range.
Ṡ_routine Ṡ_routine 1 L/h working in hot environments OBSERVED-REPLICATED Nutritional Needs in Hot Environments (NCBI NBK236240) Field measurement outside 0.5–2 L/h.
Ṡ_max Ṡ_max 3.7 L/h n = 1 — Alberto Salazar, 1984 Olympic Marathon. A single reported record maximum: one runner, one race, uncorroborated OBSERVED-SINGLE (single report) primary: Armstrong, Hubbard, Jones & Daniels 1986, Physician Sportsmed 14(3):73–81; secondary route: NBK236240 A higher documented rate; any replication at n > 1.
E_L(300) E_L(300) 2.87 × 10⁻²¹ J per bit erased k_B T ln2, T = 300 K MODELED (computed) Landauer 1961, IBM J Res Dev 5(3):183–191; k_B exact per SI 2019 Arithmetic error.
E_L(310) E_L(310) 2.97 × 10⁻²¹ J per bit erased k_B T ln2, T = 310 K (body) MODELED (computed) as above Arithmetic error.
P_brain P_brain ~20 W human brain, ~2% body mass, ~20% of resting metabolism OBSERVED-REPLICATED (not primary-sourced in this pass) standard cerebral-metabolism references (Clarke & Sokoloff, Basic Neurochemistry) Fetch a primary calorimetric/CMR source; a value outside ~15–25 W.
E_bit,syn E_bit,syn ~10⁴ ATP per bit chemical synapse, blowfly retina OBSERVED-REPLICATED Laughlin, de Ruyter van Steveninck & Anderson 1998, Nat Neurosci 1(1):36–41 Independent measurement >10× off under stated conditions.
E_bit,graded E_bit,graded 10⁶–10⁷ ATP per bit graded signals in photoreceptor/interneuron, or spike coding OBSERVED-REPLICATED as above As above.
c²/h c²/h 1.36 × 10⁵⁰ bits·s⁻¹·kg⁻¹ Bremermann's limit MODELED (computed) Bremermann 1962, Self-Organizing Systems; c, h exact per SI 2019 Arithmetic error.
m_brain m_brain ~1.4 kg adult human brain mass, as used for the Bremermann figure OBSERVED-REPLICATED (standard reference; not primary-sourced in this pass) standard anatomical references Fetch a primary source; a value outside ~1.2–1.5 kg.
v_myel v_myel up to 150 m/s myelinated axons OBSERVED-REPLICATED Purves et al. 2001, Neuroscience 2nd ed., Sinauer Measurement outside stated range.
v_unmyel v_unmyel 0.5–10 m/s unmyelinated axons OBSERVED-REPLICATED as above As above.
τ_callosal τ_callosal 3–9 ms fastest cross-brain time, 95th-percentile axons, 14 anthropoid primates over a 97-fold brain-mass range OBSERVED-REPLICATED Phillips et al. 2015, Proc R Soc B 282:20151535 (corr. 282:20152620) An anthropoid with 95th-percentile cross-brain time outside 3–9 ms.
τ_callosal,median τ_callosal,median 11–38 ms cross-brain conduction time for median axon diameters, same 14 anthropoid primates — the bulk of interhemispheric traffic OBSERVED-REPLICATED Phillips et al. 2015 An anthropoid with median cross-brain time outside 11–38 ms.
d_callosal d_callosal majority < 1 µm callosal myelinated axons, all 14 species OBSERVED-REPLICATED Phillips et al. 2015 A species with majority > 1 µm.
N_neur,human N_neur,human 86 × 10⁹ (cortex ~16 × 10⁹) neurons human brain, isotropic fractionator OBSERVED-REPLICATED Azevedo et al. 2009, J Comp Neurol 513(5):532–541 Independent count >2× off.
N_neur,elephant N_neur,elephant 257 × 10⁹ total; 251 × 10⁹ (97.5%) in cerebellum; cortex 5.6 × 10⁹ neurons African elephant OBSERVED-REPLICATED Herculano-Houzel et al. 2014, Front Neuroanat 8:46 Independent count >2× off.
N_neocort,pilot whale N_neocort,pilot whale ~37.2 × 10⁹ neocortical neurons long-finned pilot whale — ~2× the human figure OBSERVED-REPLICATED Mortensen et al. 2014, Front Neuroanat 8:132 Independent count >2× off.
N_FlyWire N_FlyWire 139,255 neurons; >50 × 10⁶ synapses complete adult Drosophila connectome, proofread OBSERVED-REPLICATED Dorkenwald et al. 2024, Nature Independent reconstruction differing materially.
B_sensory B_sensory ~10⁹ bits/s human sensory data-gathering MODELED Zheng & Meister 2025, Neuron 113(2) Recompute from receptor counts and rates.
B_behaviour B_behaviour ~10 bits/s human behavioural throughput — a literature synthesis, not a new experiment: same paper and same method as B_sensory MODELED-CONTESTED Zheng & Meister 2025, Neuron 113(2) (Perspective); contested by Sauerbrei & Pruszynski 2025, Nat Neurosci ('The brain works at more than 10 bits per second') Carry both. A measure reconciling cognitive throughput and motor control under one accounting.

sha256 e443dca15319ea0b — of the original file, so what was ingested stays checkable.